The leaf
Insertion of Leaves : The point at which a
leaf is attached to the stem or branch is called its insertion. Leaves are variously inserted on the stem and branches and accordingly different terms are given. Leaves are called radical when they arise at or below the surface of the ground and thus spring apparently from the root, but really from a shortened stem ; Examples-Taraxacum officinale (Compositae), Primula vulgaris (Primulaceae), species of Xyris (Xyridaceae) etc. The leaves which arise from the main stem with distinct internodes between them are called cauline, and those from the branches ramal.
Phyllotaxy : Phyllotaxy is the mode of arrangement of leaves on the stem and the branch. The main principle of this arrangement of leaves on the stem and the branch is to avoid shading one another so that each and every leaf may get maximum amount of sunlight to perform their normal function, Eames and MacDaniels (1947) have also stated that “leaf arrangement on the stem, or phyllotaxy, and variations in size and shape are doubtless in part correlated with the exposure of the photosynthetic surface to light.” This arrangement of leaves in a definite manner may also bear an intimate structural relation to the vascular, skeleton system of the axis.
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Different types of phyllotaxy |
Phyllotaxy is of three principal types viz : ( i ) spiral or alternate or acyclic, (ii) opposite and (iii) whorled 0r verticillate. Opposite and whorled or verticillate types are commonly called cyclic type of phyllotaxy.
1. SPIRAL OR ALTERNATE PHYLLOTAXY (i.e. acyclic type )In this type, leaves are arranged at each node in a spiral manner all round the stem. If an imaginary Spiral line is drawn winding round the stem, it will pass through consecutive nodes. Hence it is termed as spiral phyllotaxy. This imaginary spiral line is known as genetic spiral. When imaginary vertical lines are drawn from upwards and downwards, it is found that all the leaves of the plant lie in a fixed number on these vertical lines ; these vertical straight lines passing through leaves are known as orthostichies (singular, orthostichy). The distance formed upon the spiral of the stem between two successive leaves is called divergence. The angle sebtended by the divergence at the centre is known as angular divergence. The angular divergence can be found out by the following formula:
The angular divergence=number of circles or circles/no of leaves or orthostichies of a circle i.e.360°.
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